"script ... is missing!" error in player when PlayableDirector used in AssetBundle, but not player (code stripping)

Reproduction steps:

1. Open the attached project from Google Drive (Ritter.zip)
2. Open the custom build UI (Build menu)
3. Make sure IL2CPP forced and Full Asset Bundles selected
4. Press Build -> Build And Run
5. Open the player logs

Expected result: Player Build runs without errors
Actual result: Errors in player.log: The referenced script (Unknown) on this Behaviour is missing!
The referenced script on this Behaviour (Game Object '<null>') is missing!

Note: the project is built using IL2CPP (This error isn't present in the mono version of the project)

Reproducible with: Timeline 1.4.8 (2020.3.32f1), 1.5.2 (2020.3.32f1), 1.7.1 (2020.3.32f1)

Could not test with: 2019.4.37: Custom packages errors - The type or namespace name 'ScriptedImporter' could not be found (are you missing a using directive or an assembly reference?) etc.
2021.2.17f1: Custom packages errors: multiple scripts console errors: error CS0518: Predefined type 'System.Index' is not defined or imported, error CS0656: Missing compiler required member 'System.Index..ctor'
2022.1.0b13, 2022.2.0a8: RenderTarget already allocated error (Custom package error)