Crash on Transform::RebuildTransformHierarchy when saving Scene after exiting the Play Mode

Reproduction steps:
1. Open the attached “BugRepro” project
2. Open the “Assets/Scenes/EngravingAnimation.unity” Scene
3. Enter the Play Mode
4. Open any Script file
5. Made some changes in the file and save it (ctrl+s)
6. Exit the Play Mode
7. Save the Scene (ctrl+s)
6. Observe the Crash

Reproducible with: 6000.3.4f1

Reproducible on: Windows 11
Not reproducible on: No other environment tested

Note: Not reproduced by CQA

First few lines of the stack trace:
{{0x00007FF8A5EEE713 (Unity) Transform::RebuildTransformHierarchy }}
{{0x00007FF8A6CBD61A (Unity) SynchronizeMeshRenderersAndMaterials }}
{{0x00007FF8A6CBD54A (Unity) AssetHotreload }}
0x00007FF8A7C1A0E2 (Unity) AssetHotreload
{{0x00007FF8A7C8E673 (Unity) HotreloadV2 }}