Sprite stops tracking associated bones in subsequent Play Mode runs when "Reload Domain" is disabled

Reproduction steps:

1. Open the attached “Bugtest.zip“ project
2. Open the “Assets/Scenes/SampleScene.unity“ Scene
3. Enter the Play Mode
4. Observe the animation in the Scene View
5. Exit/Enter the Play Mode
6. Observe the animation in the Scene View

Expected result: The Sprite is moving, and the animation plays as expected
Actual result: The Sprite is frozen and doesn’t follow associated bones

Reproducible with: 2023.1.20f1, 2023.2.4f1, 2023.3.0b1
Not reproducible with: 2021.3.33f1, 2022.3.16f1

Reproducible on: M1 MacOS 14.2
Not reproducible on: No other environment tested

Note: Sometimes it requires a few Enter/Exit Play Mode cycles for the issue to appear